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4-HNE represents a highly reactive lipid peroxidation byproduct that induces cellular damage by interacting with proteins, DNA, and other cellular components through iron- and oxygen-mediated chain-breaking reactions (7)

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Judging from the fact that γ-GCS and GS proteins do not associate and that their gene expression is controlled differently [141], it is reasonable to consider that in certain situations, γ-GCS acts dominantly compared with GS and produces γ-glutamyl peptides other than glutathione